A quite important parameter is missing from that study.
"studies were performed with the same type of lithium-ion cell"
Which type of lithium-ion cell?
Some of the most common are:
Lithium Cobalt Oxide. ...
Lithium Manganese Oxide. ...
Lithium Iron Phosphate (LFP) ...
Lithium Nickel Manganese Cobalt Oxide (NMC) ...
Lithium Nickel Cobalt Aluminum Oxide. ...
Lithium Titanate.
They each have different characteristics. Some, like LFP (which is mentioned in the study), are not as sensitive to low or high state of charge as others. LFP also have higher number of charge/discharge cycles than most, while holding similar amount of capacity. LFP have low energy density, and are therefore rarely used in electric vehicles, so are not really relevant to "how to best charge your Zero battery". They are ideal for some other uses.
Cycle count is irrelevant if it's lacking information on how big/long the cycle is. It depends on the exact battery chemistry, and how long you store the battery at each extreme.
Your link contains these statements:
"High battery temperatures increase calendar aging by accelerating parasitic side-reactions. Thus, the battery temperature should be kept low during the long nonoperating periods. " Page 144. So not "the better for storage, both in hot and cold conditions".
"Keeping the SoC at a low or medium level and lowering the battery temperature minimizes calendar aging." Page 141. So not "the lower the SOC, the better for storage".
Empty = low SOC.
"Aging of Lithium-Ion Batteries in Electric Vehicles" figure 28 clearly shows the lower the SOC, the better for storage, both in hot and cold conditions.
Ideal storage SOC at high temperature would be empty.